Mothra is an ancient moth goddess that resides on the tropical paradise called "Infant Island". She serves as the guardian of a Earth and nature, as well as a group of natives on the island and has.two tiny twin priestesses called "The Shobijin", who serve as her translators to humanity.

Godzilla was awakened by nuclear testing in 1954 and declared war on humanity as revenge. The Earth Defense Force was formed to combat Godzilla and the numerous other kaiju that appeared over the years, but was never able to defeat Godzilla.